Research Professor Emeritus
- B.S., Molecular Biology University of Washington, 1973.
- M.S., Fisheries University of Washington, 1975.
- Ph.D., Genetics The Pennsylvania State University, 1980.
Conservation biology; population genetics of natural populations, threatened and endangered species; genetic mapping of quantitative trait loci; genomic manipulation of commercially important aquatic species; effect of environmental stressors on natural gene pools.